Nongame and Rare Species Program:
Federal and State Listed Invertebrate Species

Crustaceans	State Status	Federal Status (Listed)
Diminutive Amphipod Gammarus hyalleloides NatureServe Explorer profile		Endangered
Pecos Amphipod Gammarus pecos NatureServe Explorer profile		Endangered
Peck's Cave Amphipod Stygobromus pecki NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Endangered	Endangered
Insects	State Status	Federal Status (Listed)
Coffin Cave Mold Beetle Batrisodes texanus NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Helotes Mold Beetle Batrisodes venyivi NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Comal Springs Riffle Beetle Heterelmis comalensis NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Endangered	Endangered
American Burying Beetle Nicrophorus americanus NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
A Ground Beetle Rhadine exilis NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
A Ground Beetle Rhadine infernalis NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Tooth Cave Ground Beetle Rhadine persephone NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Comal Springs Dryopid Beetle Stygoparnus comalensis NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Endangered	Endangered
Kretschmarr Cave Mold Beetle Texamaurops reddelli NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Mollusks	State Status	Federal Status (Listed)
Pecos assiminea Snail Assiminea pecos NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Endangered	Endangered
Texas Pigtoe Fusconaia askewi NatureServe Explorer profile	Threatened
Triangle Pigtoe Fusconaia lananensis NatureServe Explorer profile	Threatened
Texas Fatmucket Lampsilis bracteata NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Threatened	Candidate
Sandbank Pocketbook Lampsilis satura NatureServe Explorer profile	Threatened
Southern Hickorynut Obovaria jacksoniana NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Threatened
Louisiana Pigtoe Pleurobema riddellii NatureServe Explorer profile	Threatened
Texas Hornshell Popenaias popeii NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Threatened	Candidate
Texas Heelsplitter Potamilus amphichaenus NatureServe Explorer profile	Threatened
Salina Mucket Potamilus metnecktayi NatureServe Explorer profile	Threatened
Diamon Tryonia Pseudotryonia adamantina NatureServe Explorer profile		Endangered
Phantom Springsnail Pyrgulopsis texana NatureServe Explorer profile		Endangered
Golden Orb Quadrula aurea NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Threatened	Candidate
Smooth Pimpleback Quadrula houstonensis NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Threatened	Candidate
False Spike Quadrula mitchelli NatureServe Explorer profile	Threatened
Texas Pimpleback Quadrula petrina NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Threatened	Candidate
Mexican Fawnsfoot Truncilla cognata NatureServe Explorer profile	Threatened
Texas Fawnsfoot Truncilla macrodon NatureServe Explorer profile U.S. Fish and Wildlife Service profile	Threatened	Candidate
Phantom Tryonia Tryonia cheatumi NatureServe Explorer profile		Endangered
Gonzales Tryonia Tryonia circumstriata NatureServe Explorer profile		Endangered
Spiders and Relatives	State Status	Federal Status (Listed)
Robber Baron Cave Meshweaver Cicurina baronia NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Madla Cave Meshweaver Cicurina madla NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Bracken Bat Cave Meshweaver Cicurina venii NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Government Canyon Bat Cave Meshweaver Cicurina vespera NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Government Canyon Bat Cave Spider Neoleptoneta microps NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Tooth Cave Spider Neoleptoneta myopica NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Tooth Cave Pseudoscorpion Tartarocreagris texana NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Cokendolpher Cave Harvestman Texella cokendolpheri NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Reddell Harvestman Texella reddelli NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered
Bone Cave Harvestman Texella reyesi NatureServe Explorer profile U.S. Fish and Wildlife Service profile		Endangered